What does the knowledge of Von Hippel-Lindau (VHL) disease and genetics reveal about related evolution concepts?

Tracking Evolution

Done by detection in people

Polymerase Chain Reaction tests

Comparative Genomic Hybridization tests

Nature of Evolution

Divergent Evolution

Requires loss of heterozygosity in chromosome 3p

Tumors developed from a germline mutation

Convergent Evolution

Different tumors becoming similar as evolution occurs

DNA Variance and Mutation

Impact

Not on number of evolving disabilities

Not on sequence of evolving disabilities

Propagation

Negative Phenotypic Manifestations

Benign and cancerous tumors

In women

Pregnancy complications

Stop reproduction

Associated with tumors in different organs

Evolution of Disease in Particular Organs

Cerebellar Hemangioblastomas

3.5-year median evolution time

Brainstem Hemangioblastomas

Progress after 7 years

Median evolution time of 1.5 years

Renal Cysts and Carcinomas

First progression after approximately 7 years

Following evolutions occur after about 1 to 2 years

May progress to clear cell renal cell carcinomas

Evolve further through additional genetic alterations

Leads to intra-tumour heterogeneity

Tumor Phylogeny

Early Truncal Events

Inactivation of the VHL gene

Mutation in chromosome 3

Some parallel evolution observed

Caused by a mutation in VHL (a tumor suppressor) gene

Leads to VHL protein abnormalities

Protein Evolution

pVHL19 is conserved more

pVHL30 is conserved less

Not Necessarily Harmful

Doesn't always stop reproduction

Would allow for mutation to be passed down

Adequate life expectancy for reproduction

Males

Average of 59.4 years

Females

Average of 48.4 years

Inheriting Germline Mutation

These might never manifest negative phenotypes

Second mutation required may not occur

Phenotypic manifestations may skip generations

Inheritance

Autosomal Dominant Inheritance Pattern

50% for a parent to pass it to their child

Different cysts

Signs of disease shown as early as 26 years old